Development of Creosote Bushes and
Clones
Recently, however, another line of evidence suggested that
individual creosote bushes were already growing at low elevations in the Mojave
Desert long before the decrease in pinyon and juniper in that region, and
perhaps even before the time that creosote bushes colonized the Chihuahu.an
Desert (as indicated by fossils of pack rat nests). This new tine of evidence
indicates that clones of creosote bush may have been continuously in residence
in the southwest Mojave Desert for nearly 12,000 years.
The latter line
of evidence is based on the development and growth rate of individual creosote
bushes and clones. Upon germination of a creosote bush seed, the first stem
grows to a height of only a few cm when a branch is formed. The main stem area
from which the first branch originates is the stem "crown" from which future
branches also originate. Later branches originate around the periphery of the
stem crown. As the stem crown grows in diameter, the center of the plant is
occupied by old branches and new branches continue to be produced around the
periphery of the stem crown. The branches grow upward a meter or so, the whole.
shrub having an obconic or somewhat rounded shape.
As growth continues,
the oldest branches gradually die and the stem crown splits or segments into
functionally separate crowns. This happens at an age of 30 to 90 years,
depending on the location. On sites with better soil moisture conditions,
growth is more rapid and segmentation occurs earlier. After segmentation
of the stern crown, the original stein and the early branches die and
eventually rot away. The connections between adjoining segments of the
Excavated plant at the time of early segmentation;
about 89 yars old. Note the relatively short, stout stem crown below the
branches; note the smaller diameter of outer (younger) branches; also note
sandy gravelly soil. |
Excavated young "seedling," with the upper parts of
branches removed, about 9 years old. Note the several branches at about the
same level and the solid, cylindrical stern crown below the. branch crotch;
this stem crown would be solid and nearly circular in cross section.
Top view of segmenting plant with tops of stems
removed. Note dead center and dead central branches, and live branches around
the periphery; plant about 49 years old but developmentally more segmented than
the plant in the above photo. |